Did Google Just Say Links In PDFs Treated As Nofollow Links?

Google's John Mueller said at the 9:43 mark into a Google hangout that for SEO purposes, links in PDF files are treated like links that have the nofollow attribute on them. But the weird thing is, Gary Illyes from Google in 2011 documented on the Google blog that those links do pass PageRank and signals. So which is it?...